About Ancestry:
When you join Ancestry, you join a human-centered company where every person’s story is important. Ancestry®, the global leader in family history, empowers journeys of personal discovery to enrich lives. With our unparalleled collection of more than 40 billion records, over 3 million subscribers and over 23 million people in our growing DNA network, customers can discover their family story and gain a new level of understanding about their lives. Over the past 40 years, we’ve built trusted relationships with millions of people who have chosen us as the platform for discovering, preserving and sharing the most important information about themselves and their families.

We are committed to our location flexible work approach, allowing you to choose to work in the nearest office, from your home, or a hybrid of both (subject to location restrictions and roles that are required to be in the office- see the full list of eligible US locations HERE). We will continue to hire and promote beyond the boundaries of our office locations, to enable broadened possibilities for employee diversity.

Together, we work every day to foster a work environment that's inclusive as well as diverse, and where our people can be themselves. Every idea and perspective is valued so that our products and services reflect the global and diverse clients we serve. 

Ancestry encourages applications from minorities, women, the disabled, protected veterans and all other qualified applicants. Passionate about dedicating your work to enriching people’s lives? Join the curious.

At Ancestry, we’re all about helping people discover, preserve, and share their family stories. If you want to join a collaborative, mission-driven team focused on delivering meaningful experiences at scale, we’re growing — and we’d love to meet you.

We’re looking for a Director of UX to lead our End-to-End Experience team — a cross-functional group focused on making the customer journey seamless across every interaction, channel, and product surface. This is a rare opportunity to shape how millions of users discover and engage with their family history through cohesive, intuitive, and deeply meaningful experiences. You’ll report to the Head of Design for the Core Product, and work closely with peers across product, design, engineering, research, content, marketing, and customer support. Your job is to ensure every step of the customer journey — from onboarding and discovery to upgrades and long-term engagement — feels thoughtfully connected and optimized.

Ancestry serves millions of customers globally. Your leadership will help unify our ecosystem of products into a seamless experience that builds trust, reinforces value, and empowers people to explore their identities and family stories. You’ll join a highly collaborative environment that values curiosity, clarity, and a deep commitment to customer understanding. We offer a flexible, hybrid culture with a strong focus on work/life balance — so you can grow your career while staying grounded in what matters most.

What you will do...

• Lead the design vision and execution for the End-to-End Experience team, ensuring cohesion and delight across the full customer journey.

• Partner across product areas and functions to eliminate friction, fill gaps, and ensure customers experience Ancestry as a rich, connected, and purpose-driven platform.

• Identify and prioritize key moments in the user journey that drive trust, retention, and discovery — and improve them through thoughtful product design and iteration.

• Collaborate closely with product, engineering, data, marketing, and customer support to ensure every touchpoint is aligned and intentional.

• Translate insights from qualitative research and quantitative data into actionable design direction, balancing creativity and clarity.

• Evangelize the customer journey internally — bringing a user-first lens to cross-org decision making and fostering a culture of customer empathy.

• Guide the creation of key design artifacts, including user flows, prototypes, wireframes, and visual systems.

• Coach and mentor a team of talented designers, helping them collaborate and grow as strategic thinkers and empathic customer advocates.

Who you are...

• Extensive experience in UX or product design, including a track record of leading design in cross-functional, collaborative environments.

• Proven ability to move between strategic thinking and hands-on execution with ease.

• Deep understanding of end-to-end customer journey mapping and experience design.

• Experience leading teams focused on onboarding, lifecycle, navigation, and engagement.

• A strong foundation in human-centered design, with the ability to bring clarity to complex or unfamiliar spaces.

• Strong communicator and influencer who can align diverse stakeholders around a shared vision.

• Passionate about creating connected, high-trust experiences that put the user first.

• Bonus: Background in consumer subscription products, digital ecosystems, or identity-driven experiences.

Helping people discover their story is at the heart of ours. Ancestry is the largest provider of family history and personal DNA testing, harnessing a powerful combination of information, science and technology to help people discover their family history and stories that were never possible before. Ancestry’s suite of products includes: AncestryDNA, AncestryProGenealogists, Fold3, Newspapers.com, Find a Grave, Archives.com, and Rootsweb. We offer excellent benefits and a competitive compensation package. For additional information, regarding our benefits and career information, please visit our website at http://ancestry.com/careers

As a signatory of the ParityPledge in Support of Women and the ParityPledge in Support of People of Color, Ancestry values pay transparency and pay equity. We are pleased to share the base salary range for this position: $172,000 - $216,000 with eligibility for bonus, equity and comprehensive benefits including health, dental and vision. The actual salary will vary by geographic region and job experience. We will share detailed compensation data for a specific location during the recruiting process. Read more about our benefits HERE.

*Note: Disclosure as required by sb19-085(8-5-20) and sb1162(1-1-23)

Additional Information:

Ancestry is an Equal Opportunity Employer that makes employment decisions without regard to race, color, religious creed, national origin, ancestry, sex, pregnancy, sexual orientation, gender, gender identity, gender expression, age, mental or physical disability, medical condition, military or veteran status, citizenship, marital status, genetic information, or any other characteristic protected by applicable law. In addition, Ancestry will provide reasonable accommodations for qualified individuals with disabilities.

All job offers are contingent on a background check screen that complies with applicable law.  For San Francisco office candidates, pursuant to the San Francisco Fair Chance Ordinance, Ancestry will consider for employment qualified applicants with arrest and conviction records.  

Ancestry is not accepting unsolicited assistance from search firms for this employment opportunity. All resumes submitted by search firms to any employee at Ancestry via-email, the Internet or in any form and/or method without a valid written search agreement in place for this position will be deemed the sole property of Ancestry. No fee will be paid in the event the candidate is hired by Ancestry as a result of the referral or through other means.